舞台是充满兽耳、兽尾、有翼族的古装、异族、战国时代，据说在服饰、固有名词等方面参考了现在生活在北海道中北部的阿伊努民族。剧情以受伤倒在路边的面罩男哈库奥罗接受素未谋面的兽耳少女艾露露的治疗为起点，经历了哈库奥罗领导人民起义反抗暴政、领导卫国斗争、解放他国受压迫民众的战争到保卫全世界大致四个阶段。大陆的历史以及哈库奥罗究竟是谁、这些异民族又是如何产生等问题在战争的扩大化中逐步得到解决。
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